From what I've heard, Bollers and Penders did not like each other very much.
Yeah pretty much. I heard a story, so dunno if it's true or not, but basically Penders was a right prick to work with. Penders expected Bollers and other writers to follow what he had laid out in the series and what he was going to lay down in upcoming story arcs, despite the fact that Penders wasn't the editor. So he wanted them to follow his work and read his scripts, but he refused to follow any of their own work if he didn't have to. And supposedly knowing this, Bollers did that storyline which resulted in a bunch of Echidna characters being killed off by using them in those Egg Grape things. Ken was supposedly really butthurt about that. And if that story is true, then it's all the more awesome.
The Egg Grapes made Eggman a psychopath. Might as well have had him built a bomb powerful enough to destroy reality itself.